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They have dreams. Someday, the dream of flying to the vast sky. Knowing how far away the sky covered by glass is. The distant future. Humanity built a mobile fortress city called "Plantation" on the desolate land and celebrated civilization. Among them, the pilot residential facility called "Mistilteinn," commonly known as "Birdcage," is where the children live. They know nothing about the outside world. They know nothing about the sky of freedom. The only mission they are told is to fight. The enemy is the enormous life form shrouded in mystery, called "Dragon." To combat an unseen enemy, the children board robots known as "FRANXX." They believe that by riding them, they are proving their existence. There was a once-called prodigy, a boy with the codename 016, named Hiro. But now, he has fallen to the bottom. He is a person who is not needed by anyone. If he doesn't board a FRANXX, he is as if he doesn't exist. In front of such a Hiro, one day, a mysterious girl called 02 appears. On her forehead, there are two vivid horns. "——I've found you, my DARLING."