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In the Sahara Desert border of West Africa, the chimpanzee David, who has ruled his group for three years, faces challenges from other males. To protect his reign, he begins to form alliances. When under siege, the indomitable king embarks on a vengeful journey to reclaim his throne. In Antarctica, thousands of emperor penguin fathers gather together to withstand the extreme cold of the endless night. They wait for the birth of new life and for their loved ones to return. On the Kenyan savannah, the lioness Cham fights to protect her pride. In the face of a great disaster, she must confront a life-or-death test. In a national park in Zimbabwe, the elderly tri-colored wolf Tete feels increasingly overwhelmed, and her control over her pack is waning. Her daughter is rising, coveting her mother's throne. In India, the tigress Rajabala hunts prey and raises four cubs all by herself, but nature never shows her any mercy.